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Leonardo also studied and drew the anatomy of many animals, dissecting cows, birds, monkeys, bears, and frogs, and comparing in his drawings their anatomical structure with that of humans. Leonardo was a modern of Botticelli, Ghirlandaio and Perugino, who have been all slightly older than he was. He would have met them at the workshop of Verrocchio or at the Platonic Academy of the Medici. Piero della Francesca had created a detailed study of viewpoint, and was the initial painter to make a scientific study of light. These research and Leon Battista Alberti’s treatise De pictura were to have a...